Oldest craniates and vertbrates in Cambrian (early Paleozoic Era); earliest gnathostomes in Silurian (middle Paleozoic Era); split between actinopterygians and sarcopterygians in Devonian (middle Paleozoic Era); oldest stegocephalians in Late Devonian Epoch; split between amphibians and amniotes in Mississippian (late Paleozoic Era).
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